Revisión actualizada de la estratigrafía y biota del Ediacárico y Terranóvico de los Montes de Alcudia-Toledo, Zona Centroibérica, España ; Updated stratigraphic framework and biota of the Ediacaran and Terreneuvian in the Alcudia-Toledo Mountains of the Central Iberian Zone, Spain

An updated stratigraphic subdivision of the Ediacaran and Terreneuvian in the Alcudia valley and the Toledo Mountains, Central Iberian Zone, is documented here. The Lower Alcudian-Domo Extremeño Supergroup is subdivided, from bottom to top, into the conformable Guadiana (La Coronada and Sta. Ma de Zújar formations) and Campanario (Botija, Monroy and Orellana formations) groups. The supergroup contains biostratigraphically non-significant bacterial acritarchs and dubious fossils, but its Ediacaran age is constrained by detrital zircon analyses. This siliciclastic sedimentary package is unconformably overlain by either the siliciclastic-carbonate Ibor Group (Castañar, Villarta and Arrocampo formations) or the siliciclastic Cíjara Formation. The fossil content of the former group includes Sabelliditids, Vendotaenids, macrophytes, bioaccumulations and reefs rich in Cloudina, Sinotubulites, Protolagena and stromatolites and thrombolites, and a wide diversity of ichnofossils; whereas the Cíjara Formation has yielded bacteria attributed to Bavlinella and Palaeogomphosphaeria, and simple trace fossils (Gordia, Helminthoidichnites and treptichnids). The Ediacaran-Cambrian boundary, based on ichnofossils, lies at the base of the Arrocampo Formation and the uppermost part of the Cíjara Formation. In the Alcudia valley, the Ibor Group is subdivided into another three-fold, lithologically equivalent subdivision, known as the Tamujar, Hinojosas (rich in ichnofossils, such as treptichnids, Monomorphichnus, Psammichnites and Taphrelminthopsis) and Cabezarrubias (including Bergaueria and Planolites) formations. Another unconformity, intra-Fortunian in age, marks the tops of the Ibor Group and the Cíjara Formation, wich are subsequently overlain by the San Lorenzo Formation and some megabreccia beds, respectively. Overlying the slope-related Fuentes, Recibido el 4 de junio de 2019; Aceptado el 21 de agosto de 2019; Publicado online el 12 de noviembre de 2019 Citation / Cómo citar este artículo: Álvaro J.J. et al. (2019). Updated stratigraphic framework and biota of the Ediacaran and Terreneuvian in the Alcudia-Toledo Mountains, Central Iberian Zone, Spain. Estudios Geológicos 75(2): e093. https://doi.org/10.3989/ egeol.43620.548. Copyright: © 2019 CSIC. This is an open-access article distributed under the terms of the Creative Commons Attribution-Non Commercial (by-nc) Spain 4.0 License. 2 Álvaro J.J. et al. Estudios Geológicos, 75(2), julio-diciembre 2019, 75(2), e093, ISSN-L: 0367-0449. https://doi.org/10.3989/egeol.43620.548 Introduction The Iberian Massif of the western Iberian Peninsula was subdivided by Lotze (1945) into several Variscan tectonostratigraphic domains or “zones”, which are currently used in pre-Variscan and Variscan palaeogeographic and geodynamic reconstructions (Fig. 1A). The Central Iberian Zone was formally defined by Julivert et al. (1972) by grouping together Lotze’s Galician-Castilian and East Lusitanian-Alcudian Zones. Both sub-domains differ in the relative abundance of Variscan syn-orogenic granitoid bodies associated with high-grade metamorphic rocks (more abundant in the former), but the contact between the two sub-domains is not sharp but gradational. In the NW segment of the Central Iberian Zone, several allochthonous complexes (Galicia-Trás-os-Montes Zone; Farias et al., 1987) are superposed, which will not be considered below, such as the Cabo Ortegal and Ordenes complexes and the Malpica-Tuy Band in Galicia (Spain), and the Bragança and Morais complexes in Trás-osMontes (Portugal; Ribeiro et al., 1990) (Fig. 1A). One of the stratigraphic peculiarities of the Central Iberian Zone is the presence of a Furongian gap, in some cases scouring deeper and eroding the entire Cambrian and part of the uppermost Ediacaran. As a result, the widespread ‘Purple Series’ and Armorican Quartzite (Lower Ordovician) occur unconformably overlying an inherited palaeorelief composed of Ediacaran-‘lower Cambrian’ strata. The Furongian uplift and subsequent denudation is related to the so-called Toledanian Membrillar and Navalpino megrabreccia beds, the Terreneuvian-Cambrian Series 2, heterolithic Pusa Formation is subdivided into thre members, the middle one characterized by the record of phosphate ore deposits (e.g., the Fontanarejo Bed). The formation is shale dominated and has yielded the body-fossil Beltanelliformis and numerous ichnofossils (Monomorphicnus and Treptichnus; lower member), sponge spicules associated with thrombolites (middle member), and Scenella, trilobites, archaeocyaths and ichnofossils (Dactyloidites, Psammichnites, Rusophycus and Teichichnus; upper member). Where the megabreccia beds are absent, the Ibor Group is unconformably overlain by the San Lorenzo and Fuentepizarra formations, the latter containing phosphatic limestone interbeds rich in skeletonized microfossils, such as Anabarella, which represents the Fortunian-Cambrian Stage 2 boundary interval.


Introduction
The Iberian Massif of the western Iberian Peninsula was subdivided by Lotze (1945) into several Variscan tectonostratigraphic domains or "zones", which are currently used in pre-Variscan and Variscan palaeogeographic and geodynamic reconstructions (Fig. 1A). The Central Iberian Zone was formally defined by Julivert et al. (1972) by grouping together Lotze's Galician-Castilian and East Lusitanian-Alcudian Zones. Both sub-domains differ in the relative abundance of Variscan syn-orogenic granitoid bodies associated with high-grade metamorphic rocks (more abundant in the former), but the contact between the two sub-domains is not sharp but gradational. In the NW segment of the Central Iberian Zone, several allochthonous complexes (Galicia-Trás-os-Montes Zone; Farias et al., 1987) are superposed, which will not be considered below, such as the Cabo Ortegal and Ordenes complexes and the Malpica-Tuy Band in Galicia (Spain), and the Bragança and Morais complexes in Trás-os- Montes (Portugal;Ribeiro et al., 1990) (Fig. 1A).
One of the stratigraphic peculiarities of the Central Iberian Zone is the presence of a Furongian gap, in some cases scouring deeper and eroding the entire Cambrian and part of the uppermost Ediacaran. As a result, the widespread 'Purple Series' and Armorican Quartzite (Lower Ordovician) occur unconformably overlying an inherited palaeorelief composed of Ediacaran-'lower Cambrian' strata. The Furongian uplift and subsequent denudation is related to the so-called Toledanian Membrillar and Navalpino megrabreccia beds, the Terreneuvian-Cambrian Series 2, heterolithic Pusa Formation is subdivided into thre members, the middle one characterized by the record of phosphate ore deposits (e.g., the Fontanarejo Bed). The formation is shale dominated and has yielded the body-fossil Beltanelliformis and numerous ichnofossils (Monomorphicnus and Treptichnus; lower member), sponge spicules associated with thrombolites (middle member), and Scenella, trilobites, archaeocyaths and ichnofossils (Dactyloidites, Psammichnites, Rusophycus and Teichichnus; upper member). Where the megabreccia beds are absent, the Ibor Group is unconformably overlain by the San Lorenzo and Fuentepizarra formations, the latter containing phosphatic limestone interbeds rich in skeletonized microfossils, such as Anabarella, which represents the Fortunian-Cambrian Stage 2 boundary interval.
During the onset of the Cadomian orogeny, the cratonward side of the back-arc basin displayed a northeastward increasing depth and availability of accommodation space (Quesada, 1991;Eguíluz et al., 1995Eguíluz et al., , 2000, then becoming a retro-arc foreland basin (Quesada, 2019a, b). Denudation of the arc can be geochronologically controlled by the input of exotic clasts (such as black chert lydites) and detrital zircon derived from the neighbouring uplifting arc (Pereira et al., 2012a, b;Sánchez-García et al., 2019). By ca. 580 Ma, the back-arc basin started to close, antithetic to the main subduction system (i.e. with the Ossa-Morena Zone as upper plate), as indicated by significant emplacement of the calc-alkaline arc-related rocks of the Malcocinado Group and related plutons in the central and (mainly) northern parts of the Ossa-Morena Zone. This gave rise to a secondary arc in this region, erupted onto/intruded into deformed, pre-orogenic to synorogenic rocks of the Serie Negra, and which extended until ca. 535 Ma (Quesada, 2019a, b).
Closure of the back-arc basin led to progressive collision of the arc with the margin of Gondwana in the range of 570-535 Ma. An oblique regime is suggested by both the large diachroneity of the collision process and the different nature of the basement across the suture (Quesada, 2019a, b). After the erosion of the Cadomian Arc, palaeocurrents changed from the present-day SW (the setting of the arc) to the NE, and the detrital zircon and clasts changed of source, from the SW Cadomian arc to the NE Cantabro-Ebroan Source Land ; Estudios Geológicos, 75(2), julio-diciembre 2019, 75(2), e093, ISSN-L: 0367-0449. https://doi.org/10.3989/egeol.43620.548 Quesada, 1991Quesada, , 1996San José et al., 1992;Rodríguez Alonso et al., 1995Eguíluz et al., 2000;Valladares et al., 2000Valladares et al., , 2002.
The suture zone, bringing the Ossa-Morena Zone onto the Central Iberian Zone, as indicated by shedding of Ossa-Morena lithologies as clasts into syn-orogenic deposits in the Central-Iberian part of the back-arc basin (Pieren Pidal, 2000), was reactivated several times (during Cambrian rifting and during the subsequent Variscan orogeny), being presently exposed within the broad Badajoz-Córdoba shear Zone and probably beneath the Obejo-Valsequillo Domain (Quesada & Dallmeyer, 1994).
In the Ossa-Morena part of the retro-arc basin, the relatively abrupt change from Cadomian subduction (latest evidence at ca. 533 Ma), with evidence of two pulses of Cadomian deformation and coeval high-grade syntectonic metamorphism related to accretion in an orogenic wedge complex, to Cambrian rifting (earliest at ca. 530 Ma; Bandrés et al., 2002Bandrés et al., , 2004Sánchez-García et al., 2019) is correlative with the three-fold geodynamic evolution recorded in the Central Iberian Zone, only loosely constrained in time due to the scarcity of radiometric data, from (i) back-arc (passive-margin) to (ii) back-arc (retroarc) foreland, and (iii) rift conditions. Three geodynamic models are at present proposed to explain this Ediacaran-Terreneuvian arc-to-rift turnover in the Iberian margin of West Gondwana, which are not necessary exclusive, involving: (i) the oblique collision of a mid-ocean ridge with the trench located at the outer Gondwana active margin, which would have progressively transformed the former subduction margin into a transcurrent one; (ii) a variant of the previous model, adding a component of subduction of the mid-ocean ridge beneath the continental upper plate; and (iii) a process of slab-pull related to the onset of subduction in the Iapetus and Prototethys oceans that may have triggered slab roll-back and back-arc rifting along the previous active margin around northern Gondwana (for a summary, see Sánchez-García et al., 2019).
In the Cadomian retro-arc basin preserved in the Central Iberian Zone, two major geodynamic episodes are recognized: (i) a late Cadomian folding event, unrelated to both schistosity and metamorphism (e.g., Apalategui et al., 2009), and stratigraphically identified as the distinct gap that separates the traditional "Lower Alcudian" (ca. 580-560 Ma) and "Upper Alcudian" (ca. 550-540 Ma on the basis of detrital zircon; Talavera et al., 2015) packages that exhibit two different styles of deformation; and (ii) the late metamorphic Sardoal event that attained medium pressure/high temperature conditions at ca. 530 Ma in the vicinity of the Badajoz-Córdoba Shear Zone (Henriques et al., 2015(Henriques et al., , 2017. The latter event broadly correlates with the intra-Terreneuvian breakdown of the former retro-arc basin and the beginning of extensional conditions broadly marked by the base of the Pusa Formation.

Structural style of the Central Iberian Zone
The southwestern boundary of the Central Iberian Zone is broadly located along the Badajoz-Córdoba Shear Zone and its northeastern boundary along the Ollo de Sapo Anticline (Díez Balda et al., 1990) (Fig. 1A). The former represents the reactivated remains of an ophiolitebearing Cadomian suture (N-MORB amphibolites and dismembered serpentinite slices; Ábalos, 1990;Quesada & Dallmeyer, 1994;Arenas et al., 2018;Díez Fernández et al., 2019), and the latter a major normal Cambrian fault that episodically separated platform and basinal settings (another example is the Vivero Fault separating the Ollo de Sapo Anticline and the Lugo Dome), and played a key role during the Variscan Orogeny as major reactivated thrust systems (Martínez Catalán et al., 1992;Russo & Bechstädt, 1994;Álvaro et al., 2010).
Three main pre-Mesozoic deformation events are superposed in the Central Iberian Zone: (i) a Cadomian folding phase, (ii) the Toledanian Phase, and (iii) the Variscan deformation.
(ii) The Furongian Toledanian Phase has been commonly mistaken with the Mid-Late Ordovician Sardic Phase (e.g., Moreno et al., 1976;Díez Balda et al., 1990) reported in Sardinia, the Occitan Domain and the Eastern Pyrenees (Álvaro et al., 2016a;Pereira et al., 2018). The Toledanian Phase, geodynamically interpreted as a Estudios Geológicos, 75(2), julio-diciembre 2019, 75(2), e093, ISSN-L: 0367-0449. https://doi.org/10.3989/egeol.43620.548 result of a break-up unconformity (Sánchez-García et al., 2019), is marked by an angular discordance in the southern Central Iberian Zone, which separates variably tilted Ediacaran-to-Cambrian Series 2 packages from an overlying Ordovician succession. The involved gap includes, at least, most of the Furongian and basal Ordovician, although the erosion can incise into the entire Cambrian and part of the Ediacaran basement (Gutiérrez-Marco et al., 2002). In contrast, in the northeastern (palaeogeographically proximal to Gondwana) Iberian Massif's units, such as the northeastern Central-Iberian, West Asturian-Leonese and Cantabrian zones and their lateral prolongation into the Iberian Chains, the rift/drift turnover is transitional and conformable, despite some local gaps in the Cantabrian Zone (Aramburu et al., 2004). The 'Purple Series' and the Armorican Quartzite contain an ubiquitous conglomeratic and quartzitic succession that broadly seals the Toledanian palaeorelief and represents the return to stable platform conditions (McDougall et al., 1987;Gutiérrez-Alonso et al., 2007;Shaw et al., 2012Shaw et al., , 2014. In the Ossa-Morena Zone, the break-up unconformity is locally capped by slope-related to alluvial breccias and conglomerates, dated at ca. 489 Ma (Venta del Ciervo K-bentonite ;López Guijarro et al., 2008), which rapidly evolved into open-shelf marine conditions that prevailed until Early Devonian times.
Associated with the Toledanian Phase, a Furongian-Early Ordovician felsic-dominant magmatic episode is recognized in the Central Iberian Zone, geographically bracketed between the Urra Formation marking its southwestern boundary and the Ollo de Sapo Formation along its northern boundary, as well as the (para) autochthonous realms of the NW Iberian Variscan Massif. Both volcanosedimentary belts represent a significant igneous event, spanned between the late Furongian and the Floian (ca. 495-470 Ma). These igneous rocks are dominantly felsic and calc-alkaline exhibiting an arc-like geochemical signature that some authors have interpreted as a result of subduction (Castro et al., 2002;Fernández et al., 2008;Del Greco et al., 2016, and references therein), whereas others argue that they were generated by melting of a subduction-related Neoproterozoic crust being its geochemical signature inherited (Díez Montes et al., 2010;García-Arias et al., 2018;Sánchez-García et al., 2019). The interplay of zircon source provenances and retrodeformation of the Ibero-Armorican Arc and Alpine superposition is providing conditions amenable to everlasting discussions on palaeogeographic interpretations (for a review, see Murphy et al., 2016). The putative association of the Toledanian unconformity and the emplacement of calc-alkaline volcanics are interpreted, in the Marão Anticline and the Amêndoa-Carvoeiro Synform of the Central Iberian Zone, as the record of compressive/transpressive tectonics (Romão et al., 2005). These authors have described cleavage-bearing folds with steep axial planes at high angles to Variscan structures, which have been interpreted as transient compression and dextral strike-slip features along the Central Iberian/Ossa-Morena zone boundary. Despite the diachronism of the Furongian Toledanian event and the Sardic intra-Ordovician phase, Romão & Ribeiro (1993), Romão et al. (2005Romão et al. ( , 2013 and Amaral et al. (2014) have interpreted the Toledanian and Sardic events as the migration, in time, of a single geodynamic regime. However, according to Hammann et al. (1982), Ribeiro et al. (1990), Quesada (1990) and Gutiérrez-Marco et al. (2002), among others, the Toledanian and Sardic Phases are separated in time. Despite the exception yielded by the Marão Anticline and the Amêndoa-Carvoeiro Synform, the Toledanian and Sardic phases are related to neither metamorphic nor cleavage features, two features characteristic of the Variscan deformation.
(iii) The Variscan deformation was responsible for the origin of two different tectonic domains in the Central Iberian Zone, the Domain of Recumbent Folds to the NE and the Domain of Vertical Folds to the SW (Díez Balda et al., 1990), roughly coinciding with the so-called Ollo de Sapo and Schist-Greywacke domains, respectively (Martínez Catalán et al., 2004). The latter, on which this paper is focused, is characterized by geographical landscapes exhibiting an Appalachian geomorphological style composed of narrow synclines topped by the 'Purple Series' and the Armorican Quartzite, and broad antiforms exhibiting Ediacaran-Cambrian successions in their cores. The overall Variscan structure of the Central Iberian Zone is the overprinting result of three distinct deformation events plus a somewhat later activity related to the subvertical shear zones or to faults (Díez Balda et al., 1990;Díez Fernández & Pereira, 2016;Dias da Silva et al., 2017).
Among the numerous anticlines of the Central Iberian Zone, for practical reasons, five Variscan tectonostratigraphic units with distinct lithological features will be described below: the Extremenian Anticlinorium, the Abenójar-Tirteafura Dome and the Alcudia, Valdelacasa and Ibor-Navalpino anticlines.

Historical background on stratigraphic terminology
The Ediacaran System of the southwestern Central Iberian Zone is composed of a thick and relatively monotonous succession of deep-water siliciclastic rocks (Pieren Pidal, 2000), which change laterally into the heterolithic sandstone/shale alternations of the Beiras Group (Medina Estudios Geológicos,75(2), julio-diciembre 2019, 75(2), e093, ISSN-L: 0367-0449. https://doi.org/10.3989/egeol. 43620.548 et al., 1998;Pereira et al., 2012b;Meireles et al., 2013). The monotonous lithologies of the Spanish side, combined with relatively scattered outcrop and later tectonic overprint, has complicated the elucidation in detail of the stratigraphic successions. Symptomatic of these problems, the relative order of lithostratigraphic units has been an endless subject of discussion (e.g., Pardo Alonso & Santamaría Casanovas, 1992). A greater lithological diversity is developed in shallower-water sedimentary rocks crossing the Ediacaran-Cambrian transition, but, even there, many uncertainties remain in the stratigraphic correlation throughout the Central Iberian Zone. As a result, a plethora of lithostratigraphic terms is available in the Spanish regional bibliography, differing between regions but also with competing schemes being developed within the same region. Much of this nomenclature was developed without formal definitions and published in PhD theses and extended abstracts. Below we provide both a brief historical background and an attempt at unifying a common lithostratigraphic terminology.
In the 1980s, and due to the geostrategic interest in the global phosphate rock market, the Spanish administration instigated the HESPERICA Project, led by the Spanish Geological Survey (IGME) and MAYASA Co., to estimate the phosphate ore reservoirs in central Spain.
The study area comprised about 54,000 km 2 of surface throughout the Central Iberian Zone. Five sectors received special attention: Fuenteaguinaldo (Salamanca province), Robledo del Mazo (Toledo) and Abenójar, Horcajo de los Montes and Fontanarejo (Ciudad Real). The latter yielded an estimate of ore resources close to 5,800,000 tons with an averaged content of about 22.7 wt. % P 2 O 5 . Some results were published in meeting proceedings and geological maps, such as Álvarez-Nava & Robles Casas (1988), , Calvet Alonso & Salas (1988), Pardo Alonso & Robles Casas (1988), Nozal Martín et al. (1988a and Ortega Gironés et al. (1988); for a historical reappraisal, see López Díaz (1994) (Fig. 3). These contributions followed a common stratigraphic sketch. In the Valdelacasa Anticline, the "Valdelacasa Group" included, from bottom to top, the Fuentes Olistostrome, the Pusa Shales, the Azorejo Sandstone and the Navalucillos Limestone formations, the two latter then considered as Cambrian due to the presence of chronostratigraphically significant ichnofossils in the former and archaeocyaths in the latter (e.g., San José et al., 1974;Gil Cid et al., 1976;Perejón et al., 1976Perejón et al., , 1981Zamarreño et al., 1976). Underlying the "Valdelacasa Group", they coined the term "Domo Extremeño" for the stratigraphically lowermost beds cropping out particularly in the areas of Las Hurdes, the Extremenian Anticlinorium and the Alcudian valley. In the Valdelacasa Anticline, they included rocks attributed to the "Estomiza and Cubilar" formations. In this scheme, the Estomiza Formation was envisaged to be in faulted contact with overlying rocks and should represent the lowermost stratigraphic part. This part of the succession, also attributed to the Cíjara Formation in other studies (Palacios Medrano, 1989), bears simple trace fossils and is clearly earlier than any rocks attributed to the "Lower Alcudian" in the Alcudia Anticline. A carbonate-bearing succession overlying the Domo Estudios Geológicos, 75(2), julio-diciembre 2019, 75(2), e093, ISSN-L: 0367-0449. https://doi.org/10.3989/egeol.43620.548 Extremeño Group was distinguished as the Ibor or Ibor-Navalpino groups, which is particularly well represented in the Navalpino and Ibor anticlines Nozal Martín et al., 1988a, b, c). The Ibor-Navalpino Group was subdivided, following San José (1984), into several lithostratigraphic subunits: the La Antigua Conglomerate, Los Parrales Shale, Villarta Limestone and Tamujosillo Shale and Greywacke formations; in the surroundings of Torilejo village, two subunits were added: the Valdehornos Rhythmite and the Torilejo Conglomerate and Shale (Fig. 3).
The stratigraphic subdivision of the Navalpino Anticline was proposed by San José (1984), who distinguished the "Lower Alcudian", "Upper Alcudian" and "Pusian" units, the two latter separated by the Navalpino Olistostrome. In the stratigraphic synthesis of the Central Iberian Zone, San José et al. (1990) maintained the triad "Lower Alcudian", "Upper Alcudian" and "Pusian", but considered them as megasequences obviating their former litho-and chronostratigraphic significance.
Another source of stratigraphic nomenclature was developed by the joint collaboration of the universities of Extremadura (Badajoz, Spain) and Uppsala (Sweden). Palacios Medrano (1989), Vidal et al. (1994a, b) and Palacios & Vidal (1996), among others, subdivided the Domo Extremeño Group into the Estenilla and Cíjara formations, maintained the Ibor Group in the Navalpino-Ibor anticlines, and proposed the Río Huso Group to include the Fuentes Olistostrome and the Pusa Shales.
The stratigraphic subdivision proposed below intends to offer a solution for such a mixture of litho-and chronostratigraphic and megasequence terminology, maintaining the lithostratigraphic terms that have not suffered from excessive conceptual changes.

Stratigraphic framework and sedimentary environments
Based on the above-reported historical revision, the Ediacaran-Terreneuvian can be subdivided in the Alcudia valley and the Toledo Mountains into several lithostratigraphic units bounded by major gaps, which are, from bottom to top: (i) the Lower Alcudian-Domo Extremeño Supergroup, (ii) the Ibor Group and its laterally correlative Cíjara Formation, the former group capped by (iii) the Fuentes, Membrillar and Navalpino megabreccia beds and (iv) the Pusa Formation, whereas the Cíjara Formation is overlain by (v) the San Lorenzo and Fuentepizarra formations. Following the rules of the International Stratigraphic Guide (Salvador, 1994;Murphy & Salvador, 1998), the stratigraphic framework proposed below highlights stratigraphic packages of similar lithological composition, separated by regional unconformities or major hiatuses (Figs. 4-5).

Lower Alcudian-Domo Extremeño Supergroup (new)
The Lower Alcudian-Domo Extremeño Supergroup represents both (i) the "Lower Alcudian" Group, Stage and megasequence of Ovtracht & Tamain (1970), Herranz et al. (1977) andSan José et al. (1990); and (ii) the lower part (underlying the Cadomian gap) of the widely used "Domo Extremeño" Group of Álvarez Nava et al. (1988). Both terms have been widely used and their conceptual modifications in time are not too significant. We propose the Alcudia valley as type area for the supergroup. Its base is not identified and its thickness is broadly estimated to be more than 6 km. The supergroup mainly consists of monotonous shales and greywackes with subsidiary sandstone (lower part) and includes clast-and matrixsupported conglomerate and breccia interbeds (upper part), and scattered centimetre-scale carbonate nodules and layers. Clasts from these conglomerates and breccias are polymictic and include intraformational (mainly greywacke and shale) and exotic (e.g., lydite, quartzite, gneiss and granite) clasts, locally associated with debris derived from pegmatite and vein-quartz remains; many exotic clasts carry internal deformation fabrics.
The lower part of the Lower Alcudian-Domo Extremeño Supergroup corresponds to both the La Coronada and Sta. María de Zújar formations of García Hidalgo et al. (1993) and Pieren Pidal (2000), and to the Guadiana, Botija, Monroy and lower part of the Orellana formations sensu Palacios et al. (2010Palacios et al. ( , 2013. Due to their lateral equivalence, the Guadiana Formation should be considered as a group subdivided into the La Coronada and Sta. María de Zújar formations. The upper part of the supergroup includes the Orellana Formation of García Hidalgo et al. (1993) and Pieren Pidal (2000), which corresponds to the upper part of the "Orellana" Formation of Palacios et al. (2010Palacios et al. ( , 2013. In order to avoid the conceptual changes associated with the Orellana toponymy, we propose that the Campanario Group should be considered as the upper part of the supergroup, subdividing it into the Botija, Monroy and Orellana formations. 1A. Guadiana Group ("Guadiana Shales" sensu Herranz et al., 1977) The Guadiana Shales sensu Herranz et al. (1977), up to 3600 m thick, comprise a monotonous succession of shales and fine-to coarse-grained greywackes, locally including subsidiary pebbly shaly lenses ( Fig. 6A-E). Its type area is defined along the La Coronada and Campanario villages. The group is subdivided into the La Coronada and Sta. María de Zújar formations.
The Sta. María de Zújar Formation (Pieren Pidal, 2000), about 600 m thick, consists of massive, mediumto coarse-grained greywackes and subsidiary microconglomerates and shales, which display common T ae Bouma sequences and "disorganized facies" (a term widely used in the regional bibliography referring to mélanges rich in slumping and matrix-and clast-supported breccia deposits) reflecting progradational slope-related fans Estudios Geológicos, 75(2), julio-diciembre 2019, 75(2), e093, ISSN-L: 0367-0449. https://doi.org/10.3989/egeol.43620.548 and proximal turbidite lobes, episodically interrupted by unconsolidated event beds (Pieren Pidal, 2000). The stratotype lies along the service road that follows the Zújar River valley (GPS N38º59'00'', W5º38'30º). Fuenlabrada et al. (2016) analyzed major and trace elements, REE, and Sm-Nd isotopes of greywacke samples from this formation. Their geochemical results point to an active margin setting as the most likely source for deposition. Trace element diagrams of the Ediacaran greywackes indicate a clear affinity to a continental island arc. The basin in which these deposits were laid down was not a part of the arc: the eroded arc was the source but the depocenter was a foreland (retro-arc) basin; a characterization of the pre-Cadomian Sm-Nd isotopes from the neighbouring Ossa-Morena Zone occurs in Rojo-Pérez et al. (2019).
In the Alcudia Anticline, the presence in the supergroup of the acritarchs Orygmatosphaeridium sp. and Protosphaeridium sp. was reported by Mitrofanov & Timofeiev (1977;unpublished report cited in San José, 1984), which has been traditionally used to date the supergroup as "Riphean-early Vendian". However, these acritarchs no longer have biostratigraphic value (e.g., Vidal et al., 1994b;Liñán & Palacios, 1982). Simón (2017) reported the trace fossil Torrowangea aff. rosei and possible Ediacara-type holdfasts from "Lower Alcudian" strata of the Alcudia Anticline. However, the discoidal structure is insufficiently diagnostic to allow any confident identification, and the biogenicity of the trace fossils is uncertain. A small looping structure described from the Membrio area (Vidal et al., 1994a) within the Guadiana Formation of Palacios et al. (2013) is probably biogenic, but could be either a trace or a filamentous fossil.

1B. Campanario Group (new)
The Campanario Group conformably overlies the Sta. María de Zújar Formation. Its stratotype lies along the road from Campanario village to the Zújar River (GPS: N38º55'40'', W5º36'00''). According to Pieren Pidal (2000), the Orellana Formation, conformably ovlerying the Sta Mª de Zújar Formation, was originally subdivided into a lower monotonous part of shales and greywackes and an upper part of matrix-supported conglomerates. Subsequently, Palacios et al. (2010Palacios et al. ( , 2013 maintained the Orellana Formation for the upper part of Pieren Pidal's (2000) homonymous unit, which is retained in this revision, and distinguished the Botija Shales and the Monroy Greywackes as separate formations conformably underlying the aforementioned matrix-supported conglomerates. As a result, the Campanario Group is subdivided by us into the Botija, Monroy and Orellana formations.
The Monroy Formation , up to 150 m thick, comprises a succession of alternating shales and fine-to medium-grained greywackes. Its stratotype lies along the eastern bank of the José Mª Oriol reservoir (Tagus River), along the road that links La Perala and Cañaveral villages (GPS: N39º42'58.41'', W6º27'1.28'').
In the vicinity of the Orellana dam, a distinct package of matrix-supported breccias and conglomerates, belonging to the Orellana Formation, has been proposed by Linnemann et al. (2018) as representative of the socalled Weesenstein-Clanzschwitz-Orellana Glaciation. Based on detrital zircon, the maximum depositional age for these strata (565 ± 4 Ma) should be significantly younger than that for the Gaskiers glaciation (ca. 579 Ma). This lithostratigraphic unit begins with conglomerates and coarse sandstones, wich are followed upsection by fine-grained sandstones and shales. Diamictites suggest features indicative of a glaciomarine origin, including dropstones, rainout sediments, flat iron-shaped pebbles and facetted pebbles. Pebble size varies from a few millimetres to 20 cm . Nozal Martín et al., 1988a, b, c) The Ibor Group is mainly recognized in the Sierra de la Zarzuela and the surroundings of the Orellana dam, the Ibor, Navalpino and Alcudia anticlines, and the Abenójar-Tirteafuera domes. It is subdivided below into the Castañar, Villarta and Arrocampo formations.

Ibor Group
2A. The Castañar Formation ("Castañar Siltstones" of Álvarez Nozal Martín, 1988a, b, c;Palacios et al., 2010), up to about 400 m, consists of shales and greywackes with minor coarse-grained sandstone-toconglomerate interbeds, and carbonate nodules and thin layers. In the surroundings of Guadalupe town, the base of the formation is marked by distinct arkosic sandstones (Palacios et al., 2013). The stratotype lies along the road that links Robledollano and Castañar de Ibor (GPS: N39º37'30'', W5º25'46''). The shale interbeds contain both Vendotaenids and simple horizontal ichnofossils. Close to Navalvillar de Ibor, Hufnagel (2008) described a greater diversity of macrophytes: "club-shaped" forms were tentatively compared with Longfengshania, although substantially smaller; a spindle-shaped form was described as Salobrigia guadalupensis and a clubshaped form as Ibora sdzuyi. This site has also yielded spirally coiled forms and rare forms branching into three.
In the Sierra de la Zarzuela and the surroundings of the Orellana dam, Pieren Pidal (1990) subdivided the strata unconformably overlying the Lower Alcudian-Domo Extremeño Supergroup into several informal units, which are proposed in this study as members of the Castañar Formation. These include, from bottom to top, the Orellanita conglomerate (Fig. 7A) and the Cogolludo sandstone (Fig. 7B)  to 50 m thick, consists of amalgamated quartzarenitic conglomerates grading upward into sandstone-dominant sandstone/shale alternations, easily recognized by their variegated colours, which have been interpreted as the onset of alluvial-to-fluvial environments (Pieren Pidal et al., 1991).
2B. The Villarta Formation (San José, 1984;López Díaz, 1994) is a heterolithic succession, ranging from 100 to 250 m in thickness. In its type area, the northern shore of the Cíjara dam, close to Villarta de los Montes village, the Villarta Formation can be subdivided into three members, from bottom to top: (i) an alternation of lenticular and bedded carbonates embedded in greenish shales ( Fig. 8B-C); (ii) a conglomeratic to sandstone-dominant unit passing upward to sandy dolostones (Fig. 8F); and (iii) an alternation of shales and bedded carbonates (Fig. 8H). The stratotype of the formation is located on the northern shore of the Cíjara dam, at the La Majada del Andaluz hut (GPS: N39º13'52.86'', W39º13'52.86'').
Cloudinid-microbial buildups are recorded in the lower member of the Villarta Formation (Fig. 8B). These have yielded, after acid etching, specimens of Cloudina (C. carinata, C. hartmanae and probably C. xuanjiangpingensis), Sinotubulites (S. baimatuoensis) and Protolagena sp. (a problematic flask-shaped microfossil; Cortijo et al., 2010Cortijo et al., , 2015 (Fig. 9A-E). Vendotaenids are relatively common in the shale interbeds of the same member (Fig. 8G), whereas Sabelliditids are abundant in the shale interbeds of the middle member. Simple ichnofossils are scattered, except in the upper member, where they are relatively common.
In the Sierra de la Zarzuela and the surroundings of the Orellana dam, Pieren Pidal (1990) distinguished a heterolithic, sandstone/shale succession, overlying the abovereported Cogolludo Sandstone Member, which comprises two distinct limestone interbeds, up to 2 m thick, named by the author "Loma de la Calera" (Fig. 8A) and "Collado de la Liebre" limestone interbeds, in some cases passing laterally into centimetre to decimetre-scale nodules of carbonate embedded in a monotonous shale. The shales between both carbonate marker beds have yielded ichnofossils, such as Treptichnus isp. The lower limestone interbed grades laterally into the Talarrubias Dolostone Bed, up to 6 m thick, rich in Sabellidites and treptichnids, and represents the lower part of the Villarta Formation in the area. The limestones of the Abenójar Dome have also yielded tubular skeletal fossils including Cloudina (Vidal et al., 1994b;Zhuravlev et al., 2012;Simón, 2018).
2C. The Arrocampo Formation (new) is proposed to solve numerous structural problems, reported in previous works. Several formations have been defined in the literature seemingly overlying the Villarta Formation, but all of them are strongly affected by structural folds and faults that preclude any correct stratigraphic setting. As a result, we propose the Arrocampo Formation as a shale-dominated unit with subsidiary sandstone interbeds increasing upwards, and local presence of centimetrescale carbonate nodules parallel to stratification. This unit is up to 350 m thick, and conformably overlies the Villarta Formation. Its stratotype lies along the right bank of the Tagus River (GPS: N39º46'51.13'', W5º44'48.71''), close to the Almaraz nuclear power plant, named Arrocampo by García-Hidalgo (1985). The monoclinal outcrops of this formation are unconformably overlain by either the San Lorenzo Formation or the Lower Ordovician conglomerates and quartzites. The base of the formation contains the earliest ichnofossils assigned to Treptichnus pedum, so tentatively marking the base of the Terreneuvian.

Cíjara Formation (modified from
The Estenilla Formation, defined by Palacios Medrano (1989) in the geographically upper part of the homonymous valley, represents two units: (i) the lower "member" displays the lithology and facies associations characteristic of the above-reported Orellana Formation, and is not retained in this work; whereas (ii) the upper part comprises the shales that mark the top of the Orellana Formation and the basal part of the Cíjara Formation, in the sense proposed here. An analysis of sulphur isotopes made in the Estenilla Formation by Strauss (2002) yielded organic carbon contents between <0.1 and 1.0 wt% (n=67) and sulphide sulphur abundances between <0.1 and 5.9 wt% (n=83). Resulting S/C ratios range from 0.1 to 7.1, with a few values substantially higher. DOP values are <0.40. δ 34 S values between -17.4 and +49.3‰ (n=14) characterize pyrite between +32.4 and +49.3‰. A most prominent feature is the positive correlation between organic carbon and sulphide sulphur, and high S/C ratios pointing to overabundance of sulphur over organic carbon.
The Cíjara Formation has yielded bacterial acritarchs attributed to Bavlinella faveolata (=Sphaerocongregus variabilis Moorman, 1974) and Palaeogomphosphaeria cauriensis Palacios Medrano (1989). Ichnofossils are represented by simple horizontal forms such as Gordia marina and Helminthoidichnites (for discussion on identification, see Jensen & Palacios, 2016). Possible treptichnids have been reported from this unit (Vidal et al., 1994b;Jensen, 2003), but it remains unclear if these represent trace fossils with multiple outlets or a vertical sinusoidal movement (cf. Jensen et al., 2006).
The Cíjara Formation includes numerous sloperelated event deposits characterized by the presence of tool, bounce and flute marks, and common channelized deposits associated with slumps and olistostromes. The presence of limestone clasts, reflecting episodes of carbonate production in laterally equivalent areas, suggests the contemporaneous deposition of the Villarta Formation during the sedimentation of parts of the Cíjara Formation. The abundance of coeval fracturing and fissuring in a substrate submitted to episodic uplift, tilting and rotation of blocks is highlighted by the abundance of hydrothermal veining and reworking of their vein quartz clast counterparts.

Fuentes, Membrillar and Navalpino megabreccia beds
The Cíjara/Pusa contact is locally marked by the onset of megabreccia beds: the Fuentes Bed in the Valdelacasa Anticline (Cortázar, 1878a, b), and the Navalpino and Membrillar Beds in the Navalpino Anticline (Moreno Serrano, 1974;Moreno, 1975Moreno, , 1977aLópez Díaz, 1992. The beds, up to 200 m thick, consist of clastsupported, unsorted conglomerates and gravel sandstones that display drastic changes in thickness (Fig. 11A-D). Their blocks, which can reach 2 m in size, are composed of limestone, dolostone, sandstone, shale and conglomerate, and are crosscut by networks of quartz veins. The larger blocks are commonly subrounded, whereas the smaller ones are (sub)angular in shape.
The megabreccia beds have been previously interpreted as gravitational slides triggered by "seismoevents" (e.g., Moreno, 1975Moreno, , 1977aSantamaría Casanovas & Remacha Grau, 1994). Although these chaotic deposits have been traditionally interpreted as "olistostromes" (a term that refers to accumulation as a semifluid body by submarine gravity sliding or slumping of unconsolidated sediments), they represent indeed submarine fan and slope-apron deposits whose sub-units can show overlapping geometries.

Pusa Formation ("Pusa Shales" of
Until the 1970s, the age of the Pusa Formation was tentatively assigned to the Vendian. Brasier et al. (1979) reported the presence of Beltanelloides ("Vendian") and Chuaria ("upper Riphean-Vendian") associated with ichnofossils, such as Monomorphichnus, which suggested the onset of a superposition of distinct Vendian and Cambrian fossil remains. San José (1983) suggested placing the base of the Cambrian at an indeterminate horizon ranging from the base of the megrabreccias to the top of the Pusa Shales.
5.1. The lower member, about 1100 m thick, shows abundant slumpings, breccia levels and contorted beds at its basal part (Figs. 11F, 12A). Basal clast-and matrixsupported breccias and contorted beds, rich in abundant hydrothermal veins and clasts, grade upward into monotonous shale (Fig. 12B) with interbedded greywacke sandstone. Phosphatic crusts are locally abundant. The only body fossils known from this member are circular (now tectonically elongated) carbonaceous compressions that have been reported under various names (see above) but are now generally attributed to Beltanelliformis. (e.g., Jensen et al., 2006;Ivantsov et al., 2014) (Fig. 11C). Their stratigraphic implications will be discussed below. Trace fossils include a range of ichnotaxa (e.g. Vidal et al., 1994b;Jensen et al., 2006;Gámez Vintaned & Liñán, 2007), such as Monomorphichnus lineatus Crimes et al. (1977), Treptichnus bifurcus Miller, 1889 from the middle part of the member. Liñán et al. (1993) proposed as stratotype section for the base of the Cordub(i)an regional Stage the aforementioned disused railtrack section, close to the La Nava de Ricomalillo. Talavera et al. (2012) reported youngest concordant detrital zircon ages of 536±13 Ma and 533±17 Ma from the approximate mid-portion of this unit; additional detrital zircon ages from the lower member would help constraining the maximum depositional age of this unit and the formation of the Fuentes Bed.
5.3. The upper member, 1900-2000 m thick, is composed of shale and greywacke sandstone with minor quartz-arenite interbeds and rare carbonates. Depositional sedimentary structures are well developed and soft-sediment slumping is common. In comparison to the underlying members of the Pusa Formation, the upper member represents relative shallower deposits, which grade upward into the conformable Azorejo Formation. Body fossils from the upper part of the member occur near Robledo de Buey, where carbonate-cemented siltstone and partly silicified nodules have yielded trilobites (cf. Abadiella bourgini) and microfossils (Cupitheca, Pelagiella and unidentified hyolithids) . This level has also yielded poorly preserved archaeocyaths, amongst which A. Zhuravlev (pers. comm. in Liñán et al., 2015: p. 1130 identified Capsulocyathus sp. Ichnofossils were reported by Jensen et al. (2010), such as Dactyloidites, Psammichnites gigas (Fig. 12D), Teichichnus and the lowest recorded occurrence of Rusophycus in the Pusa Formation. Scenella sp. has been reported across the Member 2-3 transition in the area of La Nava de Ricomalillo. (Pieren & García-Hidalgo, 1999) and Fuentepizarra (new) formations These units were defined in the Alcudia Anticline and are considered as lateral equivalents of the lower and middle members of the Pusa Formation, respectively. Due to the presence of reddish channelled conglomerates and sandstones in the neighbouring Abenójar and Tirteafuera domes, as well as variegated sandstone/shale alternations capping the Arrocampo Formation in the Ibor and Navalpino anticlines, both formations are proposed as mappable units.

Major stratigraphic gaps
Two major gaps are recognized throughout the Ediacaran-Terreneuvian of the Alcudia valley and the Toledo Mountains.

Late Cadomian gap
The base of the Ibor Group and the Cíjara Formation represents an intra-late Ediacaran paraconformable to high angular discordance that separates two sedimentary packages with different styles of deformation. The Lower Alcudian-Domo Extremeño Supergroup exhibits steeply plunging cleavage-free folds. The palaeorelief formed as a result of compressive deformation is onlapped by either the Ibor Group or the Cíjara Formation. The deformation phase, named "Oretanian Phase", among others, by San José Nozal Martín et al. (1988a, b, c), was firstly recognized in the Alcudia valley (Redlin, 1955;Crespo & Rey, 1972), although assigned to different ages. At present, it is considered as representative of a "late Cadomian folding event" (Talavera et al., 2015). No metamorphism is associated with this folding event.
Another interpretation for the onset of these steeply plunging folds was proposed by Vidal et al. (1994b), who suggested development of synsedimentary contorsion, and slumping of unconsolidated, kilometre-scale flysch deposits, in a continuous turbiditic-to-basinal infill with lack of shallowing and subaerial exposure. However, this interpretation did not take into account the uplift related to the sedimentation of alluvial-to-fluvial strata onlapping an inherited palaeorelief (e.g., the Orellanita Member of the Castañar Formation in the Sierra de la Zarzuela; Pieren Pidal et al., 1991). In addition, the authors considered the unconformity that marks the top of the Lower Alcudian-Domo Extremeño Supergroup as the same than that marking the base of the Pusa Formation, which is described below.

Base of Pusa Formation
The base of the Pusa Formation is rarely (para)conformable, but represented by the onset of either (i) channelized megabreccias (e.g., Fuentes, Membrillar and Navalpino Beds), infilled with blocks reworked from underlying formations; or (ii) mélange units reworking consolidated and unconsolidated, intraformational shales and sandstones of the basal Pusa Formation. All event beds are affected by common hydrothermal veininig, whose derived counterparts occur as ubiquitous quartz vein clasts.
This intra-Terreneuvian paraconformable to angular discordant contact reflects the episodic record of extensional perturbations of the basement, leading to the erosion and redeposition of detritus derived from underlying formations, such as the Cloudina-bearing carbonate clasts of the Villarta Formation. This contact can be interpreted as the turnover from Cadomianrelated compressive geodynamics (responsible for the input of unsorted siliciclastic clasts and preservation of high-angle substrates) to extensional conditions. In the neighbouring Ossa-Morena Zone, this intra-Terreneuvian episode is represented by the beginning of rifting conditions characterized by massive emplacement of rift-affinity igneous rocks (e.g., Sánchez-García et al., 2019).

Fossil content and chronostratigraphy
Ediacaran Unquestionable Ediacara-type fossils are not known from the Central Iberian Zone although it undoubtedly contains rocks of appropriate age. The absence of such soft-bodied metazoans is likely due to the persistence of high rates of sedimentation of immature sediments in a turbiditic complex, under generally deep-water depths with clayey background sedimentation with little sediment difference for the preservation on bedding-planes. Morphologically simple trace fossils are moderately common in the Cíjara Formation. The lower part of the Ibor Group contains abundant Vendotaenids in the Ibor and Valdemanco anticlines and the Abenójar Dome. Neither the trace fossils nor the Vendotaenids allow a precise chronostratigraphic assignment within the late Ediacaran. Skeletal fossils within carbonates of the Ibor Group, in particular Cloudina, have been considered a solid evidence for a late Ediacaran age. Although this datum remains the most parsimonious interpretation, recent papers suggest the overlap of cloudinids and what is traditionally considered earliest Cambrian small shelly fossils (Yang et al., 2016;Zhu et al., 2017). The survival of Cloudina in Cambrian times cannot be discarded (Linnemann et al., 2019).

Setting of the Ediacaran-Cambrian boundary interval
In the Valdelacasa Anticle, the presence of carbonaceous Beltanelliformis (in the lower member of the Pusa Formation), a short distance stratigraphically below or slightly overlapping the appearance of distinct Cambriantype trace fossils have been traditionally used to mark the Ediacaran-Cambrian boundary interval (Brasier et al., 1979;Liñán et al., 1984Liñán et al., , 1993. This level has remained the favoured placement by some workers (e.g. Gámez Vintated & Liñán, 1996& Liñán, , 2007. However, this is placed in doubt by the occurrence of both Monomorphichnus and Treptichnus traces from a putatively stratigraphically lower level, within the lower member, in a nearby section (Vidal et al., 1994b;Jensen et al., 2010). Fossils morphologically similar to the Beltanelliforms from the Pusa Formation have been reported from the lower part of the Soltanieh Formation in Iran (e.g. Stöcklin et al., 1964) and occur in the underlying Cíjara Formation. Shahkarami et al. (2017) discussed the stratigraphic implications of the Iranian material and comparable material from South China, and concluded that they are not reliable indicators of pre-Cambrian age.
While the placement of the Ediacaran-Cambrian boundary is somewhat uncertain within the Valdelacasa Anticline, a richer association of fossils with biochronological potential occur in the Ibor Group. Skeletal fossils, in particular Cloudina, within carbonate levels of the Ibor Group, are indicative of a late Ediacaran age. The upper part of the Ibor Group has yielded sabelliditids, including probable Sabellidites cambriensis (Vidal et al., 1994b). Treptichnus pedum occurs at the basal part of the Arrocampo Formation, representing the earliest occurrence of this index-fossil in the Central Iberian Zone.

Terreneuvian
Terreneuvian skeletal fossils are rare in the Pusa and Fuentepizarra formations. They are only known from a handful of locations where they nevertheless may be abundant. The Fontanarejo Bed (middle member of the Pusa Formation) has yielded sponge spicules embedded Estudios Geológicos, 75(2), julio-diciembre 2019, 75(2), e093, ISSN-L: 0367-0449. https://doi.org/10.3989/egeol.43620.548 in thromboid substrates (Fig. 12E-F) (Perconig et al., 1986;Reitner et al., 2012). In the vicinity of La Nava the Ricomalillo, some levels close to the middle-upper member transition have yielded cap-shaped fossils similar to Scenella (Martí Mus et al., 2008) associated with scarce acritarchs (Fig. 14C-F), comprising the only know record of non-bacterial acritarchs from the Pusa Formation in the Valdelacasa Anticline. Remarkable in this context is the occurrence in the Salamanca area of process-bearing acritarchs preserved in carbonate nodules in strata correlatable with the Pusa Formation (Díez Balda & Furnier Vinas, 1981).
Despite the absence of the helcionellid Watsonella crosbyi, a chronostratigraphically significant biomarker for the future selection of the GSSP marking the Fortunian/ Second Cambrian Stage boundary, the occurrence of the helcionellid Anabarella plana Vostokova (Gubanov in Vidal et al., 1999;Gubanov & Peel, 2003) in the Fuentepizarra Formation of the Alcudia Anticline allows identification of a biomarker close to the lower occurrence (LO) of W. crosbyi. In fact, the stratigraphic range of A. plana in the Anabar Uplift of the Siberian Platform (Kouchinsky et al., 2017) allows identification of a time span including the pre-trilobite Purella cristata and Watsonella crosbyi Zones in Siberia.

Inadequate and problematic stratigraphic nomenclature
Several stratigraphic terms repeatedly used in the literature should not be used due to their ambiguous definition and conceptual modification. The "Slate and Greywacke Complex" of Carrington da Costa (1950) and Teixeira (1954Teixeira ( , 1955 was subdivided by Herranz et al. (1977) into three (mixed) litho/chronostratigraphic units, from bottom to top, the B[o]eturian (a supposed Grenvillian basement that refers to the Serie Negra, the Ediacaran basement of the neighbouring Ossa-Morena Zone; San José, 1983), the Alcudian and the Pusian. Originally, the Alcudian was considered as a broad monotonous "flysch" lithosome (Bouyx, 1970;Parga, 1970;Vegas, 1971Vegas, , 1978Tamain, 1972Tamain, , 1975Moreno Serrano, 1974), up to 10 km thick, which included a heterolithic upper part with conglomerates and limestones. This upper part was considered "molasse" deposits by Maas (1963). Herranz et al. (1977) separated the upper heterolithic part from the lower monotonous "flysch" lithosome naming them "Lower Alcudian" and "Upper Alcudian". Originally, both terms represented mixed litho-chronostratigraphic units (groups and stages) that were separated by a distinct angular discordance. Vilas et al. (1979) and San José (1983) described the "Upper Alcudian", up to 3700 m thick, as a complex succession of fluvio-tidal deposits influenced by terrigenous and mixed platforms. Due to the presence of microbial structures and imprecise ichnofossils, the "Upper Alcudian" was dated as "Riphean-Vendian" (San José et al., 1974;Brasier et al., 1979).
The term "Pusian" was coined by San José (1983,1984) as another mixed litho-chronostratigraphic lithosome that included the Precambrian-Cambrian boundary, and was sandwiched between the "olistostrome" megabreccias that unconformably overlie the "Upper Alcudian", and the sandstone packages (Azorejo Formation) that traditionally marked the base of the Cambrian due to the presence of characteristic ichnofossils, such as Astropolithon and Psammichnites (Brasier et al., 1979). The Pusian comprised ichnofossils such as Monomorphichnus and Planolites, and soft-bodied impressions such as Beltanelloides and Chuaria. Subsequently, San José et al. (1990) considered the terms "Lower Alcudian" and "Upper Alcudian-Pusian" as two megasequences displaying the lithostratigraphic significance of a supergroup. These authors still placed the base of the Cambrian in an indeterminate position within the "Pusian".

"Cordub(i)an"
The Cordubian or Corduban Stage was formally defined by Liñán et al. (1984) to include all strata of the Iberian Peninsula bearing arthropod-like ichnofossils and underlying the earliest trilobite record "in order to separate it from the Ovetian Stage that was defined using trilobite faunas" (p. 824). The authors proposed the Sierra de Córdoba as a type area for the stage and the (H)uso river section of the Central Iberian Zone (Brasier et al., 1979;Palacios Medrano, 1989) as stratotype for its base, marked by the lowest occurrence (LO) of the ichnogenus Monomorphichnus. Other (para)stratotypes were selected by Liñán et al. (1993) at Barrios de Luna (Cantabrian Zone; Crimes et al., 1977) and Concha de Artedo (West Asturian-Leonese Zone; Crimes et al., 1977) sections, where the Ediacaran/Cambrian contact is marked by an angular discordance, and in the La Rinconada section (Central Iberian Zone; Corrales et al., 1974). The authors also suggested the LO of the ichnofossil Astropolichnus hispanicus as the top of the Corduban in the Tamames Sandstone of the Central Iberian Zone, but this is in contradiction with the base of the overlying Ovetian Stage, defined at the LO of dolerolenid trilobites (Sdzuy, 1971;Liñán et al., 1993).
Subsequently, Liñán et al. (1993) and Gámez-Vintaned & Liñán (1996) summarized and completed a list of ichnofossils characteristic of the Corduban. Gámez- Vintaned & Liñán (1996) suggested that the LO of Monomorphichnus "nearly coincides" with that of "Phycodes" pedum, as a result of which they proposed the LO of the M. lineatus-T. pedum ichnofossil assemblage as the Precambrian-Cambrian boundary in Spain and as new base for the Corduban. In their figure 3, the base of the upper Corduban Substage is marked by the LO of Rusophycus, predating that of Cruziana.
New findings of shelly fossils and acritarchs somewhat completed this sketch. Palacios & Vidal (1996) and Vidal et al. (1999) reported the presence of Cloudina in the Ibor Group, and of Anabarella, Aldanella, hyolithids (circothecids and orthothecids), aff. Mongolitubulus and chancelloriids in the Pusa and Fuentepizarra formations; bigotinid trilobite and archaeocyathan moulds were also found in these shales, as a result of which, the authors suggested that part of the Pusa Formation would be Nemakit-Daldynian to mid-Tommotian in age.
Finally, Liñán et al. (2002) subdivided again the Corduban into two substages, the bases of which were selected at the LO of M. lineatus + T. pedum and Rusophycus avalonensis, respectively. However the authors dramatically modified the concept of the stage: Liñán et al. (2002) selected the indeterminate bigotinids reported by Palacios & Vidal (1996) and Vidal et al. (1999)  with the Nemakit-Daldynian and Tommotian stages of Siberia.
In addition, Liñán et al. (2002) established an ichnofossil-based zone to mark the topmost of the Ediacaran: the Torrowangea rosei Zone. A single specimen of the ichnotaxon Torrowangea aff. rosei from the Saviñán Formation of the Paracuellos Formation, Iberian Chains, was illustrated and described by Liñán & Tejero (1988); however, this specimen was subsequently reconsidered as questionable by Jensen et al. (2007: Table 1) because "the published information does not allow for a critical evaluation". The presence of T. rosei (without "aff.") in the Pusa (Liñán et al., 2002: p. 19-20) and Cíjara   fig. 9) formations of the western Toledo Mountains has not yet been supported with any illustration.
Based on the presence of T. pedum at the basal part of the underlying Arrocampo Formation, the proposal of the base of the Corduban in the Pusa Formation as the regional Ediacaran-Cambrian boundary should be abandoned. The boundary is identified, at least, at the basal part of the Arrocampo Formation, below the major unconformity that marks the base of the Pusa Formation, and below the base of the regional Corduban Stage sensu Liñán et al. (1993) and Liñán et al. (2002).

Ediacaran-Terreneuvian carbonate production, evaporite record and phosphogenesis in West Gondwana
The Villarta Formation represents a latest Ediacaran episode of semi-continuous carbonate productivity, characterized by shallowing-upward cycles, less than 8 m thick, grading from offshore clayey substrates to marlstones/ shales with carbonate nodules and, finally, a widespread variation of carbonate substrates that include: (i) shoreface (wave-influenced) substrates, ooidal shoal complexes and Cloudina-microbial and microbial reefal frameworks. In some intra-platform blocks, the Villarta Formation can be subdivided into three members, the middle one displaying progradation of conglomerate-to-litharenite shoal complexes (e.g., the Villarta type area and Castañar de Ibor in the Ibor Anticline, and exposures surrounding Abenójar in the Alcudia Anticline). The base of the overlying Arrocampo Formation, which broadly corresponds to the LO of T. pedum, marks the final drowning of carbonate productivity on palaeohorsts and blocks and the sedimentation of persistent offshore-dominated clayey substrates. Therefore, the LO of pedum is constrained by facies (so environmental) conditions.
Laterally to the Iberian margin of West Gondwana (Fig. 15), a diachronous carbonate production is reported: (i) The southwestern prolongation of the Iberian margin has recorded the final episodes of the Pan-African Orogeny. The latter predated in time the SW-European Cadomian Orogeny, which represents its lateral continuation. In the Anti-Atlas of Morocco, the end of this orogeny marks the beginning of rifting conditions and the onset of a latest Ediacaran episode of carbonate productivity (phosphatic carbonates of the Taguedit Bed, Tabia Member, Adoudou Formation) followed by a Terreneuvian establishment of intraplatform microbially dominated carbonate production throughout the horst-and-graben framework of the Cambrian Atlas Rift (Tifnout Member, Adoudou Formation; Álvaro et al., 2014a) (Fig. 16). (ii) To the palaeogeographic north-east of the Iberian margin of Gondwana, the presence of carbonate interbeds across the Ediacaran-Cambrian transition has been reported in: (i) a Fortunian phosphatic limestone bed that punctuates the lower member of the Herrería Formation in the Cantabrian Zone (Álvaro et al., 2016b); (ii) the Fortunian Codos (phosphatic limestone) Bed of the Paracuellos Group (Álvaro et al., 2016b) in the eastern Iberian Chain, which represents the lateral prolongation of the West Asturian-Leonese Zone (Álvaro et al., 2018); (iii) the Terreneuvian phosphatic limestones of the Herault Member (Marcou Formation) in the northern Montagne Noire that onlap the Cadomian Rivernous volcanosedimentary complex (Clausen & Álvaro, 2007;Devaere et al., 2013;Álvaro et al., 2014b); and (iv) the microbial carbonates of the Puig Sec Member (Pic de la Clape Formation) that onlap the Cadomian Finestrelles volcanosedimentary complex in the Eastern Pyrenees (Padel et al., 2018a, b).
Due to the current co-existence of phosphates and carbonates, phosphogenesis was necessarily associated with carbonate productivity, whereas shallow-water carbonate factories were also related to centres of evaporitic precipitation. Relics of primary and early diagenetic evaporites demonstrate that extensive evaporitic conditions were locally associated with the evolution of the latest Ediacaran-Cambrian Epoch 2 carbonate-dominated and mixed platforms of the western Gondwana margin (Fig. 16). They are preserved as pseudomorphs after evaporites, such as "chicken-wire" and enterolithic structures, lenticular to lozenge-shaped crystals of gypsum and anhydrite relics (Álvaro et al., 2000). According to Scotese & Barret (1990), among others, the latitudinal abundance of these climatically sensitive facies can be estimated by statistical techniques: carbonates have a maximum likelihood of occurring between 10° and 30° latitude, and evaporites at 25º to 35° latitude. Therefore, the existence of an Ediacaran-Cambrian Epoch 2 Southern Hemisphere arid belt is envisaged in terms of widespread evaporites in internal platforms.
The occurrence in the late Ediacaran-Cambrian Epoch 2 throughout the Moroccan-SW European margin of West Gondwana of phosphorites (Cook & Shergold, 1986;Notholt & Braiser, 1986), reefal and ooidal grainstone carbonates, and evaporites displays a parallel SW-NE-trend migration of evaporitic and phosphoritic belts (Fig. 16). This pattern points to a primary palaeolatitudinal factor controlled by the late Ediacaran-Cambrian poleward drifting of West Gondwana. The lack of climate-sensitive facies (palaeosols, carbonates, evaporites and phosphorites) in some basins was primarily controlled by orogenic processes (also recorded in a south-west/north-east trend migration), such as the Rokelide (Taoudeni Basin), Pan-African (Morocco) and Cadomian (south-western Europe) orogens. The timetransgressive migration of phosphorites, delayed in time by comparison with subtropical evaporites, should fit with a climatic control (temperate setting) for phosphogenesis (Álvaro et al., 2016b).

Conclusions
An updated stratigraphic subdivision of the Ediacaran and Terreneuvian from the Alcudia valley and the Toledo Mountains, Central Iberian Zone, is documented. The Lower Alcudian-Domo Extremeño Supergroup is subdivided, from bottom to top, into the conformable Guadiana (La Coronada and Sta. Mª de Zújar formations) and Campanario (Botija, Monroy and Orellana formations) groups. This sedimentary package is unconformably overlain by either the Ibor Group (Castañar, Villarta and Arrocampo formations) or the Cíjara Formation. The Ediacaran-Cambrian boundary, based on ichnofossils, tentatively lies at the base of the Arrocampo Formation and the uppermost part of the Cíjara Formation. Another unconformity marks the base of the overlying Pusa Formation, which is subdivided into thre members, the middle one characterized by the record of phosphate ore deposits (e.g., the Fontanarejo Bed). In the Alcudia valley, the Ibor Group is subdivided into another threefold, equivalent subdivision, the Tamujar, Hinojosas and Cabezarrubias formations, whereas the equivalent of the lower part of the Pusa Formation is recognized as the San Lorenzo and Fuentepizarra formations.
The terms "Lower Alcudian", "Upper Alcudian" and "Pusian" are inadequate due to the mixture of litho-and chronostratigraphic features and should be ruled out. The boundaries and subdivision of the "Cordub(i)an", Estudios Geológicos, 75 (2) originally proposed as the lowermost and trilobite-free stage of the Cambrian in the Iberian Peninsula, has suffered several modifications that have changed its original meaning. Its base was defined in the Pusa Formation above the erosive unconformity that marks its base. As the first occurrence of Trychophycus pedum is now "lowered" until the basal part of the Arrocampo Formation (Ibor Group), the base of the Cordub(i)an Stage (no matter which one of its successive modifications) does not mark the base of the Cambrian. Due to the co-occurrence of subtropical facies-sensitive deposists, such as phosphorites, evaporites and carbonate factories leading to reefal and ooidal shoals, a migration of climatically sensitive belts is highlighted following a SW-NE trend, from present-day Morocco to the Iberian Peninsula and the delayed occurrence in southern France, due to the counter-clockwise rotation of Gondwana across the Ediacaran-Cambrian transition. These belts include (i) a proximal evaporitic belt, (ii) a distal phosphogenic belt and (iii) a gradual migration of centres of carbonate production of reefal (microbial and microbial-shelly) ecosystems during late Ediacaran-Cambrian Epoch 2 times.