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<front>
<journal-meta>
<journal-id journal-id-type="publisher-id">EGEOL</journal-id>
<journal-title-group>
<journal-title>Estudios Geol&#x00F3;gicos</journal-title>
<abbrev-journal-title>Estud. Geol.</abbrev-journal-title>
</journal-title-group>
<issn pub-type="epub">0367-0449</issn>
<publisher>
<publisher-name>Consejo Superior de Investigaciones Cientificas</publisher-name>
</publisher>
</journal-meta>
<article-meta>
<article-id pub-id-type="publisher-id">EG201927</article-id>
<article-id pub-id-type="doi">10.3989/egeol.43586.567</article-id>
<article-categories>
<subj-group subj-group-type="heading">
<subject>Articles</subject>
</subj-group>
</article-categories>
<title-group>
<article-title>Calcimicrobial-archaeocyath-bearing clasts from marine slope deposits of the Cambrian Mount Wegener Formation, Coats Land, Shackleton Range, Antarctica</article-title>
<trans-title-group xml:lang="es">
<trans-title>Clastos con calcimicrobios y arqueociatos procedentes de dep&#x00F3;sitos marinos del talud de la Formaci&#x00F3;n c&#x00E1;mbrica del Monte Wegener, Coats Land, Cordillera de Shackleton Ant&#x00E1;rtida</trans-title>
</trans-title-group>
<alt-title alt-title-type="running-head">Calcimicrobial-archaeocyath-bearing clasts from marine slope deposits of the Cambrian Mount Wegener Formation, Coats Land, Shackleton Range, Antarctica</alt-title>
</title-group>
<contrib-group>
<contrib contrib-type="author" corresp="yes">
<name>
<surname>Rodr&#x00ED;guez-Mart&#x00ED;nez</surname>
<given-names>M.</given-names>
</name>
<xref ref-type="aff" rid="aff0001">1</xref>
</contrib>
<contrib contrib-type="author" corresp="yes">
<name>
<surname>Perej&#x00F3;n</surname>
<given-names>A.</given-names>
</name>
<xref ref-type="aff" rid="aff0001">1</xref>
</contrib>
<contrib contrib-type="author" corresp="yes">
<name>
<surname>Moreno-Eiris</surname>
<given-names>E.</given-names>
</name>
<xref ref-type="aff" rid="aff0001">1</xref>
</contrib>
<contrib contrib-type="author" corresp="yes">
<name>
<surname>Men&#x00E9;ndez</surname>
<given-names>S.</given-names>
</name>
<xref ref-type="aff" rid="aff0002">2</xref>
</contrib>
<contrib contrib-type="author">
<name>
<surname>Buggisch</surname>
<given-names>W.</given-names>
</name>
<xref ref-type="aff" rid="aff0003">3</xref>
</contrib>
</contrib-group>
<aff id="aff0001"><label>1</label>Universidad Complutense de Madrid, Departamento de Geodin&#x00E1;mica, Estratigraf&#x00ED;a y Paleontolog&#x00ED;a, Madrid, Spain. Email: <email xlink:href="martarm@geo.ucm.es">martarm@geo.ucm.es</email>, <email xlink:href="aparqueo@ucm.es">aparqueo@ucm.es</email>, <email xlink:href="eiris@geo.ucm.es">eiris@geo.ucm.es</email>; ORCID ID: <ext-link ext-link-type="uri" xlink:href="http://orcid.org/0000-0002-4363-5562">http://orcid.org/0000-0002-4363-5562</ext-link>, <ext-link ext-link-type="uri" xlink:href="http://orcid.org/0000-0002-6552-0416">http://orcid.org/0000-0002-6552-0416</ext-link>, <ext-link ext-link-type="uri" xlink:href="http://orcid.org/0000-0003-2250-4093">http://orcid.org/0000-0003-2250-4093</ext-link></aff>
<aff id="aff0002"><label>2</label>Museo Geominero, Instituto Geol&#x00F3;gico y Minero de Espa&#x00F1;a (IGME), R&#x00ED;os Rosas, 23, 28003 Madrid, Spain. Email: <email xlink:href="s.menendez@igme.es">s.menendez@igme.es</email>; ORCID ID: Silvia Men&#x00E9;ndez: <ext-link ext-link-type="uri" xlink:href="http://orcid.org/0000-0001-6074-9601">http://orcid.org/0000-0001-6074-9601</ext-link></aff>
<aff id="aff0003"><label>3</label>GeoZentrum Nordbayern. Friedrich-Alexander-University of Erlangen-N&#x00FC;rnberg (FAU). Schlossgarten 5, 91054 Erlangen, Germany</aff>
<pub-date pub-type="epub">
<day>31</day>
<month>12</month>
<year>2019</year>
</pub-date>
<pub-date pub-type="collection">
<year>2019</year>
</pub-date>
<volume>75</volume>
<issue>2</issue>
<elocation-id>10.3989/egeol.43586.567</elocation-id>
<history>
<date date-type="received">
<day>01</day>
<month>05</month>
<year>2019</year>
</date>
<date date-type="accepted">
<day>01</day>
<month>07</month>
<year>2019</year>
</date>
<date date-type="Publicado on-line">
<day>10</day>
<month>12</month>
<year>2019</year>
</date>
</history>
<permissions>
<copyright-statement>&#x00A9; 2019 CSIC</copyright-statement>
<copyright-year>2019</copyright-year>
<license license-type="open-access" xlink:href="https://creativecommons.org/licenses/by/4.0/">
<license-p>This is an open-access article distributed under the terms of the Creative Commons Attribution-Non 4.0 International License</license-p>
</license>
</permissions>
<abstract>
<title>ABSTRACT</title>
<p>The carbonate clasts from the Mount Wegener Formation provide sedimentological, diagenetic and palaeontological evidences of the destruction and resedimentation of a hidden/unknown Cambrian carbonate shallow-water record at the Coats Land region of Antarctica. This incomplete mosaic could play a key role in comparisons and biostratigraphic correlations between the Cambrian record of the Transantarctic Mountains, Ellsworth-Whitmore block and Antarctic Peninsula at the Antarctica continent. Moreover, it represents a key record in future palaeobiogeographic reconstructions of South Gondwana based on archaeocyathan assemblages.</p>
</abstract>
<trans-abstract xml:lang="es">
<title>RESUMEN</title>
<p>Los clastos carbonatados de la Formaci&#x00F3;n del Monte Wegener proporcionan evidencias sedimentol&#x00F3;gicas, diagen&#x00E9;ticas y paleontol&#x00F3;gicas de la destrucci&#x00F3;n y resedimentaci&#x00F3;n de un registro carbonatadoo c&#x00E1;mbrico oculto/no conocido en aguas someras de la regi&#x00F3;n de Coats Land en la Ant&#x00E1;rtida. Este mosaico incompleto podr&#x00ED;a jugar un papel clave en las comparaciones y correlaciones bioestratigr&#x00E1;ficas entre el registro c&#x00E1;mbrico de las Monta&#x00F1;as Transant&#x00E1;rticas, el bloque Ellsworth-Whitmore y la Pen&#x00ED;nsula Ant&#x00E1;rtica del continente ant&#x00E1;rtico. Adem&#x00E1;s, representa un registro clave en las futuras reconstrucciones paleobiogeogr&#x00E1;ficas de Gondwana meridional basadas en asociaciones de arqueociatos.</p>
</trans-abstract>
<kwd-group xml:lang="en">
<title>Keywords</title>
<kwd>Calcimicrobes</kwd>
<kwd>Archaeocyaths</kwd>
<kwd>Shackleton Range</kwd>
<kwd>Antarctica</kwd>
<kwd>Gondwana</kwd>
</kwd-group>
<kwd-group xml:lang="en">
<title>Palabras clave</title>
<kwd>Calcimicrobios</kwd>
<kwd>Arqueociatos</kwd>
<kwd>Cordillera Shackleton</kwd>
<kwd>Ant&#x00E1;rtica</kwd>
<kwd>Gondwana</kwd>
</kwd-group>
</article-meta>
</front>
<body>
<sec id="sec1">
<title>Introduction and geological setting</title>
<p>The Bruce&#x2019;s Scottish National Antarctic Expedition 1902-1904 sampled the deep sea sediments of the Weddell Sea, Antarctica. Later on, Pirie (<xref ref-type="bibr" rid="cit0024">1913</xref>) first mentioned about the presence of &#x201C;Archaeocyathinae&#x201D; from these erratic Cambrian samples. Throughout the twentieth century, different expeditions collected archaeocyaths in the dispersed Cambrian record of the Antarctica continent. The studies of the autochthonous archaeocyathan assemblages from the lower Cambrian Shackleton Limestone (Transantarctic Mountains, Hill, <xref ref-type="bibr" rid="cit0017">1964a</xref>; Debrenne &#x0026; Kruse, <xref ref-type="bibr" rid="cit0010">1986</xref>, <xref ref-type="bibr" rid="cit0011">1989</xref>), Miaolingian Nelson Limestone (Pensacola Mountains, Debrenne &#x0026; Kruse, <xref ref-type="bibr" rid="cit0011">1989</xref>; Wood <italic>et al</italic>., <xref ref-type="bibr" rid="cit0027">1992</xref>) and Furongian Minaret Formation (Ellsworth Mountains, Debrenne <italic>et al</italic>., <xref ref-type="bibr" rid="cit0012">1984</xref>) have been essential to infer the likely sources of the abundant and dispersed record of allochthonous archaeocyathan assemblages within Palaeozoic conglomerates as the Whiteout Conglomerates (Ellsworth Mountains, Debrenne, <xref ref-type="bibr" rid="cit0009">1992</xref>) to Cenozoic deposits as the Polonez Cave and Cape Melville Formations (King George Island, Morycowa <italic>et al</italic>., <xref ref-type="bibr" rid="cit0022">1982</xref>; Wrona &#x0026; Zhuravlev, <xref ref-type="bibr" rid="cit0028">1996</xref>), deep sea gravels (Weddell Sea, Gordon, <xref ref-type="bibr" rid="cit0015">1920</xref>), and glacial tills and moraines from the Whichaway Nunataks (Hill <xref ref-type="bibr" rid="cit0019">1965</xref>; Debrenne &#x0026; Kruse, <xref ref-type="bibr" rid="cit0011">1989</xref>), Shackleton Range (Stephenson Bastion and Du Toit Nunataks, H&#x00F6;fle &#x0026; Buggisch, <xref ref-type="bibr" rid="cit0020">1995</xref>) and the Transantarctic Mountains (Hill, <xref ref-type="bibr" rid="cit0018">1964b</xref>). In fact, the presence of archaeocyath--bearing clasts in other Palaeozoic records, with unknown Cambrian carbonate platforms, has allowed palaeobiogeographic comparisons between the Antarctica assemblages and others exotic archaeocyaths from South Gondwana localities (<xref ref-type="fig" rid="f0001">Fig. 1</xref>), such as the Falkland Islands (Fitzroy Tillite Formation, Stone <italic>et al</italic>., <xref ref-type="bibr" rid="cit0025">2012</xref>), South Africa (Dwyka Group, Debrenne, <xref ref-type="bibr" rid="cit0008">1975</xref>), Namibia (Dwyka Group, Perej&#x00F3;n <italic>et al</italic>., <xref ref-type="bibr" rid="cit0023">2019</xref>), and Argentina (El Jag&#x00FC;elito and Sauce Grande Formations Gonz&#x00E1;lez <italic>et al</italic>., <xref ref-type="bibr" rid="cit0013">2011</xref>, <xref ref-type="bibr" rid="cit0014">2013</xref>).</p>
<fig id="f0001">
<label>Figure 1</label>
<caption><p>A. 550-505 Ma palaeogeography of the West Gondwana assembly (simplified from Gray <italic>et al</italic>., <xref ref-type="bibr" rid="cit0016">2008</xref>), showing the occurrence of <italic>in situ</italic> Botoman archaeocyathan assemblages in Antarctica and Australia localities (1-5), modified from Perej&#x00F3;n <italic>et al</italic>. (<xref ref-type="bibr" rid="cit0023">2019</xref>). Abbreviations: Aus, Australia; K, Kalahari craton; C, Congo craton; W, West Africa; RP, R&#x00ED;o de la Plata; SF, Sao Francisco; A, Armorica; AM, Amazonia; FL, Florida, AV, Avalonia. B. Mid Cambrian-late Carboniferous of the South Gondwana palaeogeography (rotated and simplified from Boger, <xref ref-type="bibr" rid="cit0001">2011</xref>), showing the distribution of allocthonous archaeocyaths assemblages from Cambrian to Cenozoic deposits (6-12), modified from Perej&#x00F3;n <italic>et al</italic>. (<xref ref-type="bibr" rid="cit0023">2019</xref>). Australia localities: 1, Arrowie Basin; 2, Gnalta Shelf; 3, Stansbury Basin. Antarctica localities: 4, Transantarctic Mountains; 5, Pensacola Mountains; 6, Shackleton Range (this study); 7a, Whichaway Nunataks; 7b, King George Island; 8, Ellsworth Mountains. Argentina localities: 9, Falkland Islands; 12, Sierra Grande; 13, Sierras Australes. Southern Africa localities: 10, main Karoo Basin, South Africa; 11, Aranos Basin, Namibia.</p></caption>
<graphic xlink:href="EG201927-112-g001.tif" xmlns:xlink="http://www.w3.org/1999/xlink"/>
</fig>
<p>The occurrence of archaeocyath-bearing clasts as Cenozoic glacial erratics and as Cambrian marine slope conglomerates of the Cambrian Mount Wegener Formation at the Shackleton Range (<xref ref-type="fig" rid="f0002">Fig. 2</xref>) was firstly reported by Buggisch <italic>et al</italic>. (<xref ref-type="bibr" rid="cit0004">1994a</xref>), H&#x00F6;fle &#x0026; Buggisch (<xref ref-type="bibr" rid="cit0020">1995</xref>) and Buggisch &#x0026; Henjes-Kunst (<xref ref-type="bibr" rid="cit0002">1999</xref>). The Shackleton Range at the Coats Land region of Antarctica is bounded by the Slessor and Recovery glaciers which drain the East Antarctic Ice Sheet into the Weddell Sea (<xref ref-type="fig" rid="f0002">Fig. 2B</xref>). The lithostratigraphic framework of the Shackleton Range has been established after the outstanding work done by a number of geological field expeditions (see Kleinschmidt, <xref ref-type="bibr" rid="cit0021">2007</xref>) in spite of the great disconnection between outcrops and its complex tectonic structure. In the Shackleton Range, the dispersed rock outcrops mainly consist of medium-high grade amphibolite facies of the Proterozoic Shackleton Range Metamorphic Complex (Clarkson, <xref ref-type="bibr" rid="cit0006">1972</xref>) and diverse Proterozoic-Cambrian metasedimentary and metavolcanic rocks, where infracrustal and supracrustal rocks are tectonically interleaved. The remaining stratigraphic record, Ordovician red beds and molasses, Permian tillites, Jurassic tuffites and dolerites and Cenozoic tills and moraines occur more or less disconnected into the different tectonostratigraphic units developed during the Ross Orogeny (Buggisch <italic>et al</italic>. <xref ref-type="bibr" rid="cit0005">1994b</xref>; Buggisch &#x0026; Kleinschmidt, <xref ref-type="bibr" rid="cit0003">1999</xref>). In general, the Shackleton Range is interpreted as a collisional orogen as a result of the final amalgamation between East and West Gondwana during late Precambrian-Cambrian times (Tessensohn <italic>et al</italic>., <xref ref-type="bibr" rid="cit0026">1999</xref>). The Cambrian Mount Wegener Formation is forming part of the Mount Wegener Nappe (<xref ref-type="fig" rid="f0002">Fig. 2C</xref>), whose transport to the south and the low-grade metamorphic overprint were around 490 Ma as a result of Ross Orogeny (Buggisch <italic>et al</italic>., <xref ref-type="bibr" rid="cit0005">1994b</xref>).</p>
<fig id="f0002">
<label>Figure 2</label>
<caption><p>A. Location of the Shackleton Range at the Antarctica continent. The East Antarctica plate is outlined. The West Antarctic plate was built up by the amalgamation of different terranes (AP, EWB, TI, MBL). AP, Antarctic Peninsula; EWB, Ellsworth-Whitmore Block; TI, Thurston Island, MBL, Marie Byrd Land; DML, Dronning Maud Land; CL, Coats Land; TAM, Transantarctic Mountains. B. Main rock outcrops map of the Shackleton Range, showing the directions of present glaciers and their tributaries, modified from H&#x00F6;fle &#x0026; Buggisch (<xref ref-type="bibr" rid="cit0020">1995</xref>). Stars point out the location of microbial-archaeocyath-bearing clasts. 1, Read Mountains; 2, Du Toit Nunataks; 3, Stephenson Bastion. C. Detail from B, showing the partial geological map of the southern Shackleton Range, modified from Clarkson <italic>et al</italic>. (<xref ref-type="bibr" rid="cit0007">1995</xref>). OHT, Otter Highlands Thrust; MWT, Mount Wegener Thrust. D. The geological map of the Mount Wegener Formation at the southern part of the Read Mountains, modified from Buggisch &#x0026; Henjes-Kunst (<xref ref-type="bibr" rid="cit0002">1999</xref>). The occurrence of in situ microbial-archaeocyath-bearing clasts from the Mount Wegener Formation conglomerates is pointed out. MWT, Mount Wegener Thrust.</p></caption>
<graphic xlink:href="EG201927-112-g002.tif" xmlns:xlink="http://www.w3.org/1999/xlink"/>
</fig>
</sec>
<sec id="sec2">
<title>Microfacies analysis of carbonate clasts</title>
<p>The analyzed carbonate clasts from the Cambrian marine slope deposits were collected from metaconglomerates and boulders of the Mount Wegener Formation at the southern part of the Read Mountains, from the Swinnerton Ledge, Oldhamia and Trueman Terraces (<xref ref-type="fig" rid="f0002">Fig. 2D</xref>) during the EUROSHACK expedition (1993-1994). A total number of 42 thin sections have been analyzed with sedimentological, palaeontological and diagenetic purposes to reconstruct the development and destruction of calcimicrobial-archaeocyath boundstones. The carbonate clasts are accessory to primary components in polymict sandy metaconglomerates and polymict breccias from the Mount Wegener Formation which was deposited in slope to basin environments. The carbonate clasts correspond to dolostones and limestones with varied microfacies as silty/sandy mudstone, bioclastic to intraclastic wackestone/packstone, common algal and mud peloidal silty packstone/grainstone, rare ooidal and oncolitic grainstone. The bioclastic to intraclastic wackestone/packstone shows a rich fossil assemblage with trilobites, brachiopods, echinoderms, archaeocyaths and undetermined skeletal remains. Calcimicrobes belonging to <italic>Epiphyton</italic>, <italic>Renalcis</italic> and <italic>Girvanella</italic> groups and archaeocyaths are conspicuous constituents in limestone clasts, forming calcimicrobial boundstone, calcimicrobial-archaeocyathan boundstone and archaeocyathan cementstone. The initial study of pore-filling and fracture cements points out to meteoric to marine-phreatic up to burial diagenesis. The observed low grade tectonically induced fabrics (achaeocyaths tectonically elongated, mechanical twinning in calcite cements, cataclasite fabrics) were produced during the final emplacement and southwards transport of the Mount Wegener Nappe. The preliminary data from the palaeontological study shows a rich and diverse archaeocyathan assemblage with more than twenty-one genera identified. The 74% of genera belong to the Ajacicyathina Suborder, 9% to Archaeocyathina, 6% to Erismacoscinina, 5% to Loculicyathina, 4% to Dokidocyathina, and 1% to Monocyathida and Putapacyathida Orders.</p>
</sec>
<sec id="sec3">
<title>Concluding remarks</title>
<p>Prof. Dr. Werner Buggisch deceased on 6.04.2019 after a long illness. In the year 2016 we asked Professor Buggisch about the material from the Shackleton Range and about the possibility of studying the calcimicrobial-archaeocyath-bearing clasts. Professor Buggisch responded enthusiastically and sent us all the material he had. Unfortunately, although he was aware of our advances, he died before seeing the first results published. Between 1979 and 1995 he participated in four Antarctic expeditions, thus this work and future publications happen thanks to his tremendous work in Shackleton Range. A brief profile of his work can be seen on the website of the GeoZentrum Nordbayern (<ext-link ext-link-type="uri" xlink:href="https://www.gzn.nat.fau.de/2019/04/09/das-geozentrum-trauert-um-prof-dr-w-buggisch-verstorben-am-6-04-2019/">https://www.gzn.nat.fau.de/2019/04/09/das-geozentrum-trauert-um-prof-dr-w-buggisch-verstorben-am-6-04-2019/</ext-link>)</p>
</sec>
</body>
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