Trace fossils of floating reptiles from the Carnian in the Tabular Cover (S Spain)
DOI:
https://doi.org/10.3989/egeol.45687.1125Keywords:
Sedimentology, Ichnology, 3D models, Upper TriassicAbstract
Small trace fossils are recorded in the top and the bottom of a thin sandstone bed located in the upper part of the K1 unit (lower Carnian, Upper Triassic), in the Tabular Cover. In the top surface, two trace fossils have been identified: a tridactyl digitigrade elongated scratch mark with preserved triangular claws (20 mm long and 19 mm wide), with two lateral elongated (30 mm) and curved marks are located back to the margins of the tridactyl footprint, and a longitudinal furrow (185 mm long and 3 mm wide). In the bottom surface there are two autopodium impressions preserved as convex hyporeliefs, counter moulds (45 mm long, 26 mm wide, and 18 mm long, 14 mm wide). They are tridactyl, elongated scratch marks, located very close and forming an angle of 48º. Trace fossils are interpreted as produced on the bottom by small tetrapods when floating in a low energy shallow environment. In the top surface correspond to the right hindlimb and the tail. In the bottom surface the scratch marks are more elongated and may be related to pes and manus. The potential authors are uncertain. This is a new example of swimming trace fossil for the Carnian deposits of the southern and eastern Iberia.
Downloads
References
AGISOFT. Agisoft Metashape: https://www.agisoft.com/ (last accessed January 2025).
Arche, A. & López-Gómez, J. (2014). The Carnian Pluvial Event in Western Europe: New data from Ibe¬rian and correlation with the Western Neotethys and Eastern North America - NW Africa regions. Earth-Science Reviews, 128: 196-231. https://doi.org/10.1016/j.earscirev.2013.10.012
Berrocal-Casero, M., Pérez-Valera, J. A., Dos Santos, V. F. & Segura, M. (2022). Nothosaur tracks from the Middle Triassic of Spain suggest a global distribution for distinctive swim track assemblages. Lethaia, 55(1), 1-11. https://doi.org/10.18261/let.55.1.6
Besems, R. E. (1981). Aspects of Middle and Late Triassic palynology. 1. Palynostratigraphical data from the Chiclana de Segura Formation of the Linares-Alcaraz region (Se Spain) and correlation with palynological assemblages from the Iberian Peninsula. Review of Palaeobotany and Palynology, 32, 257-273. https://doi.org/10.1016/0034-6667(81)90007-5
CloudCompare (2025). CloudCompare [GPL software] Stereo: http://www.cloudcompare.org (last ac¬cessed January 2025).
Dal Corso, J., Bernardi, M., Sun, Y., Song, H., Seyfullah, L. J., Preto, N., Gianolla, P., Ruffell, A., Kustatscher, E., Roghi, G., Merico, A., Hohn, S., Schmidt, A. R., Marzoli, A., Newton, R.J ., Wignall, P. B. & Benton, M. J. (2020). Extinction and dawn of the modern world in the Carnian (Late Triassic). Science Advances, 6, eaba0099. https://doi.org/10.1126/sciadv.aba0099 PMid:32938682 PMCid:PMC7494334
De Torres, T. (1990). Primeros resultados de unas dataciones palinológicas del Keuper de la Rama Cas¬tellana de la Cordillera Ibérica, Prebético y Subbético frontal. In: F. Ortí & J. Salvany (Eds.), Formaciones evaporíticas de la Cuenca del Ebro y cadenas periféricas de Levante (pp. 219- 223). ENRESA.
Farlow, J.O., Robinson, N.J., Turner, M.L., Black, J. & Gastesy, S.M. (2018). Footfall pattern of a bottom-walk¬ing crocodile (Crocodylus acutus). Palaios, 33, 406-413. https://doi.org/10.2110/palo.2018.037
Fernández, J. (1977). La sedimentación triásica en el borde Sur-Este de la Meseta. [Doctoral Thesis dis¬sertation, Universidad de Granada]. DIGIBUG: Repositorio Institucional de la Universidad de Granada. https://digibug.ugr.es/handle/10481/84072
Fernández, J., Santos-Francés, F. & Linares, J. (1976). La sedimentación triásica entre Terrinches y Al¬badalejo (Ciudad Real). Estudios Geológicos, 32, 249-259.
Luhmann, T., Robson, S., Kyle, S. & Boehm, J. (2020). Close-Range Photogrammetry and 3D Imaging. (3rd Edition). De Gruyter, Berlín. https://doi.org/10.1515/9783110607253
McCrea, R.T., Pemberton, S.G. & Currie, P.J. (2004). New ichnotaxa of mammal and reptile tracks from the Upper Paleocene of Alberta. Ichnos, 11, 323-339. https://doi.org/10.1080/10420940490442313
Milner, A. R. C., Lockley, M. G. & Kirkland, J. I. (2006). A large collection of well preserved theropod dinosaur swim tracks from the Lower Jurassic of Moenave Formation, St. George, Utah. New Mexico Museum of Nature and Science Bulletin, 37, 315-328.
Navarro, O. & Moratalla, J. J. (2023). El Barrancazo: a new locality of possible semi-aquatic turtle tracks from the Upper Triassic of Cortes de Pallás (Eastern Iberia). Journal of Iberian Geology, 49, 169- 188. https://doi.org/10.1007/s41513-023-00212-y
QGIS (2025). QGIS Geographic Information System, 3.34.11-Prizren. Open Source Geospatial Founda¬tion. http://qgis.org (last accessed February 2025)
Reolid, J. & Reolid, M. (2017). Traces of floating archosaurs: an interpretation of the enigmatic trace fossils from the Triassic of the Tabular Cover of Southern Spain. Ichnos, 24, 222-233. https://doi.org/10.1080/10420940.2016.1265524
Reolid, M., Márquez-Aliaga, A., Belinchón, M., García-Forner, A., Villena, J. & Martínez-Pérez, C. (2018). Ichnological evidence of semi-aquatic locomotion in early turtles from eastern Iberian during the Carnian Humid Episode (Late Triassic). Palaeogeography, Palaeoclimatology, Palaeoecology, 490, 450-461. https://doi.org/10.1016/j.palaeo.2017.11.025
Reolid, J., Cardenal, J., Reolid, M. & Mata, E. (2020). 3D Imaging of southernmost Triassic archosaur footprints from Europe (Southern Spain). Journal of Iberian Geology, 46, 145-161. https://doi.org/10.1007/s41513-020-00125-0
Xing, L., Lockley, M. G., Klein, H., Romilio, A. & Persons IV, W. S. (2023). A possible turtle swim track from the late Triassic of the Sichuan Basin, China. Historical Biology, 36(4), 820-824. https://doi.org/10.1080/08912963.2023.2185885
Downloads
Published
How to Cite
Issue
Section
License
Copyright (c) 2025 Consejo Superior de Investigaciones Científicas (CSIC)

This work is licensed under a Creative Commons Attribution 4.0 International License.
© CSIC. Manuscripts published in both the print and online versions of this journal are the property of the Consejo Superior de Investigaciones Científicas, and quoting this source is a requirement for any partial or full reproduction.
All contents of this electronic edition, except where otherwise noted, are distributed under a Creative Commons Attribution 4.0 International (CC BY 4.0) licence. You may read the basic information and the legal text of the licence. The indication of the CC BY 4.0 licence must be expressly stated in this way when necessary.
Self-archiving in repositories, personal webpages or similar, of any version other than the final version of the work produced by the publisher, is not allowed.
Funding data
Ministerio de Ciencia e Innovación
Grant numbers PID2023-147440OB-C21
Universidad de Jaén
Grant numbers TEP-213 y RNM-200







